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With the brief summer now passed, the animals of the Highlands are in for the long haul. Only the toughest will survive the Highlands' darkest and most overwhelming season. Reindeer, ptmarigan and mountain hare are well adapted to these extreme conditions while red squirrels and crested tits are pushed to the limit. Atlantic salmon return to spawn in the rivers in which they were born while on the Orkney island of Copinsay, grey seals haul onto the beaches to breed and give birth.

This film was first broadcast: 25 May 2016

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