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Built in response to the development of the Thameslink scheme, a large scale infrastructure project to connect northern and southern London by heavy rail, the Class 319s would be created as the backbone of this new service, and would ply its merry trade across the vital Bedford to Brighton run for a dutiful 30 years.

However, through a slight of hand by Network Southeast, more Class 319s than were needed to operate Thameslink were procured, and thus beyond its crosscapital duties, the 319 would find a variety of work initially across southern England, and later to the far reaches of the northwest, with conversions of 319s into bimode and hydrogen fuel units seeing these machines continue to work across vast swathes of the network.

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References:
Key Model World (and their respective sources)
Kent Rail (and their respective sources)
Wikipedia (and its respective references)